List of Chromosome Count Reports - Stevia

Probable Base Number for Genus x = 11, 12

NOTE: apomixis and associated chromosomal irregularities are common in Stevia

NOTE 2: the number x=17 has also been claimed for the genus, but Watanabe et al. 2001 were unable to verify this, and provide an explanation for why this claim is probably not valid.

References are at the bottom

S. alatipes 2n=22 ref Watanabe et al. 2001

S. ambryolepis 2n=24 + 1-3B ref Watanabe et al. 2001

S. alternifolia 2n=11 II + 0-1B, 2n=22 ref Watanabe et al. 2007

S. anadenotricha n=12 Grashoff et al. 1972; Watanabe et al. 2001

S. andina n= ca 22 ref Turner et al. 1967

S. aschenborniana 2n=24, 24 + 1-2B ref Strother & Panero 2001; Watanabe et al. 2001

S. bangii 2n=44 + 0-5B ref Watanabe et al. 2007

S. berlandieri n=12; 2n=24 ref King et al. 1995; Grashoff et al. 1972; Pinkava & Keil 1977

S. boliviensis n=11 ref Robinson et al. 1989

S. camachensis 2n=22 ref Watanabe et al. 2007

S. caracasana n=33 I; 2n=34I ref Robinson et al. 1989; Grashoff et al. 1972; 2n=22 + 1-2B, 33, 33 + 0-2B Watanabe et al. 2001

S. chilapensis 2n=22 ref Watanabe et al. 2001

S. cinerascens 2n=22 ref Watanabe et al. 2007

S. clausseni 2n=22 ref Watanabe et al. 2007

S. clinopodioides 2n=48 refs Watanabe et al. 2001; 2n=36 + 0-4B Watanabe et al. 2007

S. coahuilensis 2n=33 ref Watanabe et al. 2001; Watanabe et al. 2007

S. commixta 2n=22 ref Dematteis et al. 2007; Watanabe et al. 2007

S. connata 2n=22, 22+ 0-2B, 33, 44 ref Watanabe et al. 2001; Watanabe et al. 1995

S. constricta 2n=22 + 4-8B, 444 ref Watanabe et al. 2001

S. crassifolia 2n=33 ref Watanabe et al. 2001

S. decumbens 2n=24, 36 ref Watanabe et al. 2001

S. decussata 2n=22 ref Watanabe et al. 2007

S. deltoidea 2n=33, 33+0-3 B, 44, 44 + 0-6B, 55 + 0-5B ref Watanabe et al. 2001

S. deltoidea x nepetifolia 2n=33 ref Watanabe et al. 2001

S. dictyophylla 2n=24 ref Watanabe et al. 2001

S. ecatepecana 2n=12 II ref Watanabe et al. 2001

S. elatior n=11, ca 23, ca 33I ref King et al. 1995; Grashoff et al. 1972

S. elatior var. elatior 2n=33, 44 ref Watanabe et al. 2001

S. elatior var. podophylla 2n=33 ref Watanabe et al. 2001

S. ephemera 2n=8 ref Watanabe et al. 2001

S. eupatoria 2n=24, 34 I, 36, 48 ref Grashoff et al. 1972; Watanabe et al. 1995

S. filodecaballoana 2n=22 ref Watanabe et al. 2001

S. glandulosa 2n=24 ref Watanabe et al. 2001

S. hintoniorum 2n=22, 33, 33 + 1B, 44, 44 + 1B ref Watanabe et al. 2001; Watanabe et al. 2007

S. hypomalaca 2n=22 + 0-2B, 22 + 1B ref Watanabe et al. 2001

S. iltisiana 2n=33, 44 ref Watanabe et al. 2001

S. incanescens 2n=40-47 I ref Grashoff et al. 1972

S. incognita 2n=33, 33 + 0-2B, 33 + 2B, 33 + 1-3B, 44 + 0-2B ref Watanabe et al. 2001; Watanabe et al. 2007

S. isomeca 2n=35II, 33, 33+ 1-3B, 44 + 1-4B, 44 + 4B ref Sundberg et al. 1986; Watanabe et al. 2001

S. jalicensis n=12; 2n=24 ref Turner & Flyr 1966; Watanabe et al. 2001

S. jorullensis 2n=33, 34I ref Grashoff et al. 1972; 2n=24, 36, 48 Watanabe et al. 2001; 2n=36, 36 + 0-1B Watanabe et al. 2007

S. karwinskiana 2n=22, 33 ref Watanabe et al. 2001

S. latifolia 2n=33, 44, 55 ref Watanabe et al. 2001

S. lehmanii 2n=22 + 1B ref Watanabe et al. 2001

S. lemmonii 2n=24 ref Watanabe et al. 2001

S. leptophylla 2n=11 II + 4B; 22 + 0-1B, 11 II ref Frederico et al. 1996; Watanabe et al. 2007

S. liebmannii 2n=33 ref Watanabe et al. 2001

S. lita 2n=5 II ref Strother & Panero 2001; 2n=10 II, 20 Watanabe et al. 2001

S. lucida n=12, 2n=24 ref Grashoff et al. 1972; Powell & King 1969; King et al. 1995; Watanabe et al. 2001

S. lundiana 2n=22, 22 + 1B ref Watanabe et al. 2007

S. macvaughii 2n=24 ref Watanabe et al. 2001

S. mascotensis 2n=44 ref Watanabe et al. 2001

S. mercedensis n=11 ref Robinson et al. 1989

S. mexicana 2n=11 II, 22 ref Watanabe et al. 2001

S. micradenia n=33 I ref Turner & Flyr 1966; Watanabe et al. 2001

S. micrantha 2n=22 ref Grashoff et al. 1972; 2n=24 Watanabe et al. 2001

S. microchaeta 2n=24 ref Watanabe et al. 1995; Grashoff et al. 1972

S. mitopoda 2n=24 ref Watanabe et al. 2007

S. monardifolia 2n=22, 33, 34I ref Watanabe et al. 2001; Watanabe et al. 1995; Grashoff et al. 1972

S. myriadenia 2n=11II, 22 ref Frederico et al. 1996; Watanabe et al. 2007

S. myricoides 2n=12II, 24 ref Watanabe et al. 2001

S. nelsonii 2n=24 ref Watanabe et al. 2001

S. nepetifolia 2n=33, 33 + 1B, 33 + 3B, 44 + 1-4B ref Watanabe et al. 2007

S. oaxacana 2n=24 ref Watanabe et al. 2001

S. obovata n=11 ref Robinson et al. 1989

S. occidentalis 2n=20 ref Watanabe et al. 2001

S. oligophylla 2n=22, 33 ref Soejima et al. 2001

S. ophryophylla 2n=11 II, 22 ref Frederico et al. 1996; Watanabe et al. 2007

S. organensis 2n=22 ref Watanabe et al. 2007

S. origanoides 2n= 34 I; 43 I ref Grashoff et al. 1972

S. ovata n=34I, 36I, ca 60I, 2n=33, 2n=34I, 22 + 0-2B, 33 + 0-2B, 33 + 3-8 B, 44 + 2B ref King et al. 1976; Watanabe et al. 1995; Grashoff et al. 1972; Watanabe et al. 2007

S. palmeri 2n=44 ref Watanabe et al. 2001

S. parvifolia 2n=11 II, 22 ref Watanabe et al. 2007

S. perfoliata 2n=22 ref Watanabe et al. 2001

S. phlebophylla 2n=24 ref Watanabe et al. 2001

S. pilosa 2n=ca 34 ref Watanabe et al. 1995; Grashoff et al. 1972; n=11, 2n=22, 22+0-4B, 33, 44 Watanabe et al. 2001; 2n=33, 33 + 0-1B, 44 Watanabe et al. 2007

S. plummerae n= 46 I; 34, 34I, 42I ref Turner et al. 1967; Grashoff et al. 1972; 2n=22+1-2B, 44 Watanabe et al. 2001

S. polycephala n=12, 2n=12II, 24 ref King et al. 1976; Robinson et al. 1989; Watanabe et al. 2001

S. porphyrea 2n= 34 I ref Grashoff et al. 1972; 2n=22, 33, 44, 55 Watanabe et al. 2001; 2n=44 Watanabe et al. 2007

S. potosina 2n=33, 33 + 0-1B ref Watanabe et al. 2001; Watanabe et al. 2007

S. purpusii 2n= 46 I ref Grashoff et al. 1972; 2n=36 Watanabe et al. 2007

S. rebaudiana 2n=11 II ref Frederico et al. 1996

S. reticulata 2n=22, 33 ref Watanabe et al. 2001

S. revoluta 2n=24 ref Watanabe et al. 2001

S. rhombifolia n= 46 I ref Turner et al. 1967

S. rosei 2n=44 ref Watanabe et al. 2001

S. rotundifolia 2n=33 + 0-2B ref Watanabe et al. 2001

S. rzedowskii 2n=12II, 24 ref Watanabe et al. 2001

S. salicifolia 2n=24 (+0 or 1B) ref Watanabe et al. 1995; Grashoff et al. 1972; Pinkava & Keil 1977

S. samaipatensis n=11 ref Robinson et al. 1989

S. santacruzensis n=11 + 1 I ref Robinson et al. 1989

S. sarensis n=11 ref Robinson et al. 1989

S. saturiaeafolia 2n=11 II ref Sundberg et al. 1986; Dematteis et al. 2007

S. scabrella 2n=24 ref Watanabe et al. 2001

S. seemannii 2n=33, 33+0-1B, 44, 55+2B ref Watanabe et al. 2001

S. seemannioides 2n=22 ref Watanabe et al. 2001

S. seleriana 2n=12II ref Watanabe et al. 2001

S. selloi 2n=11 II, 22 ref Frederico et al. 1996; Watanabe et al. 2007

S. serrata n=ca 27, ca 33I, ca54I; 2n=33 (+0-2B), 22, 33I, 34I, 31-37I, 42-48I, 44, 44 + 1-2B, 55, 66 ref King et al. 1976; Powell & King 1969; Watanabe et al. 1995; Grashoff et al. 1972; Watanabe et al. 2001; Watanabe et al. 2007

S. serrata x porphyrea 2n=44 ref Watanabe et al. 2001

S. setifera n=11, 2n=22 ref Robinson et al. 1989; Watanabe et al. 2007

S. stenophylla n=12 ref Turner & Flyr 1966

S. stolonifera 2n=22, 22+2B, 44 ref Watanabe et al. 2001

S. suaveolens 2n=33 (+0-2B) ref Watanabe et al. 1995

S. subpubescens n=12, ca 12; 2n=24 ref Turner & Flyr 1966; King et al. 1995; Grashoff et al. 1972; Watanabe et al. 2001

S. tenuis 2n=22 ref Watanabe et al. 2007

S. tephra 2n= 24 II or 48 I ref Grashoff et al. 1972; 2n=33 + 1-2B, 33 + 3B, 33 + 0-1B, 33+3-4B Watanabe et al. 2001, Watanabe et al. 2007

S. tephrophylla n=12, ca 12 ref King et al. 1995

S. tomentosa 2n=33 ref Watanabe et al. 1995; Watanabe et al. 2001; 2n=33 + 2B, 33+3B, 44+2-4B Watanabe et al. 2007

S. triangularis 2n=24 ref Watanabe et al. 2001

S. trifida n=11, 2n=22 ref King et al. 1995; Watanabe et al. 2001

S. triflora 2n= 34-36 I ref Grashoff et al. 1972

S. tunariensis n=11 ref Robinson et al. 1989

S. urticifolia n=11 ref Robinson et al. 1989

S. vaga n=12 ref Turner et al. 1979

S. velutinella 2n=24 ref Watanabe et al. 2001

S. veronicae 2n=11 II, 22 ref Watanabe et al. 2007

S. viejoana 2n=11 II, 22 ref Watanabe et al. 2001

S. viscida 2n=ca 34; 22, 22I, 33I, 44I ref Strother & Panero 2001; Grashoff et al. 1972; Turner & Flyr 1966; Watanabe et al. 2001

S. yaconensis n=11 ref Robinson et al. 1989

S. zacatecana 2n=12II, 24 ref Watanabe et al. 2001

S. zephyrantha 2n=24 ref Watanabe et al. 2001

References

Dematteis, M., J. Molero, M. B. Angulo, and A. M. Rovira. 2007. Chromosome studies on some Asteraceae from South America. Botanical Journal of the Linnean Society 153: 221-230.

Frederico, A. P., P. M. Ruas, M. A. Marin-Morales, C. F. Ruas, and J. N. Nakajima. 1996. Chromosome studies in some Stevia Cav. (Compositae) species from southern Brazil. Brazilian Journal of Genetics 19: 605-609.

Grashoff, J. L., M. W. Bierner, and D. K. Northington. 1972. Chromosome numbers in North and Central American Compositae. Brittonia 24: 379-394.

King, R. M., D. W. Kyhos, A. M. Powell, P. H. Raven, and H. Robinson. 1976. Chromosome numbers in Compositae. XIII. Eupatorieae. Annals of the Missouri Botanical Garden 63: 862-888.

Pinkava, D. J. and D. J. Keil. 1977. Chromosome counts of Compositae from the United States and Mexico. American Journal of Botany 64: 680-686.

Powell, A. M. and R. M. King. 1969. Chromosome numbers in the Compositae: Colombian species. American Journal of Botany 56: 116-121.

Robinson, H., A. M. Powell, G. D. Carr, R. M. King, and J. F. Weedin. 1989. Chromosome numbers in Compositae, XVI: Eupatorieae II. Annals of the Missouri Botanical Garden 76: 1004-1011.

Soejima, A. Yahara, T. and Watanabe K. 2001. Distribution and variation of sexual and agamospermous populations of Stevia from Mexico. Plant Species Biology 16: 91-105.

Strother, J. L. and J. L. Panero. 2001. Chromosome studies: Mexican Compositae. American Journal of Botany 88: 499-502.

Sundberg, S., C. P. Cowan, and B. L. Turner. 1986. Chromosome counts of Latin American Compositae. American Journal of Botany 73: 33-38.

Turner, B. L. and D. Flyr. 1966. Chromosome numbers in the Compositae. X. North American species. American Journal of Botany 53: 24-33.

Turner, B. L., J. Bacon, L. Urbatsch and B. Simpson. 1979. Chromosome numbers in South American Compositae. American Journal of Botany 66: 173-178.

Turner, B. L. A. M. Powell, and J. Cuatrecasas. 1967. Chromosome numbers in Compositae. XI. Peruvian species. Annals of the Missouri Botanical Garden 54: 172-177.

Watanabe, K., R. M. King, T. Yahara, M. Ito, J. Yokoyama, T. Suzuki, and D. J. Crawford. 1995. Chromosomal cytology and evolution in Eupatorieae (Asteraceae). Annals of the Missouri Botanical Garden 82: 581-592.

Watanabe, K., T. Yahara, A. Soejima, and M. Ito. 2001. Mexican species of the genus Stevia (Eupatorieae, Asteraceae): Chromosome numbers and geographical distribution. Plant Species Biology 16: 49-68.

Watanabe, K., T. Yahara, G. Hashimoto, Y. Nagatani, A. Soejima, T. Kawahara, and M. Nakazawa. 2007. Chromosome numbers and karyotypes in Asteraceae. Annals of the Missouri Botanical Garden 94: 643-654.